terrestrial 226 soil 173 aquatic 143 lentic 81 paludal 79 wetland 79 very low temperature 59 permafrost 58 forest 53 ice 44 low temperature 34 host-associated 24 anthropogenic 22 rhizosphere 21 plant host 21 fresh water 16 low salinity 16 acidic 14 mine 12 contaminated 12 tundra 11 grassland 11 medium temperature 9 agriculture 6 lotic 5 spring 3 insect host 3 hot spring 3 animal host 3 cropland 3 fluvial 2 lacustrine 2 sediment 2 benthic 2 subsurface 1 high temperature 1 intestine 1 marine 1 digestive tract 1