terrestrial 24 aquatic 23 soil 19 lentic 17 low salinity 15 fresh water 15 benthic 11 sediment 11 lacustrine 10 grassland 8 wetland 7 paludal 6 very low temperature 6 anthropogenic 6 subsurface 5 groundwater 4 forest 3 plant host 2 host-associated 2 rhizosphere 2 littoral 1 high temperature 1 marine 1 spring 1 mangrove 1 low temperature 1 lotic 1 hot spring 1